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FidoNet consists of approximately 10,000 systems world-wide which comprise a network which exchanges mail and files via Modems using a proprietary protocol.

They are connected for the purposes of exchanging E-Mail to the Internet thru a series of gateway systems which interact with the Internet via UUCP with cooperating UNIX-based smart-hosts which act as their MX-receivers.

More info on http://www.fidonet.org/